`J_F = L_p * "S" *(( P_C - P_T ) - sigma *( Pi_C - Pi_T ))`

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The Starling equation describes the role of hydrostatic and osmotic forces in the movement of fluid across capillary endothelial cells.

Where:

• `J_F`=  Transcapillary flux
• `P_C`= Capillary hydrostatic pressure ,
• `P_T` = Tissue hydrostatic pressure 
• `pi_C`= Capillary oncotic pressure
• `pi_T`= Tissue oncotic pressure
• `L_p`= Hydraulic conductivity
• `sigma`= Plasma protein reflection coefficient
• S = capillary surface area for exchange